© Ryan Peterson

Kamchatka Steelhead Project

Kamchatka Steelhead Project

Author: Lori Howk

Abbreviated Bibliography

From the Wild Salmon Center and Moscow State University. All articles published in Voprosy Ikhtiologii are available in the translated version, Journal of Ichthyology, widely available in the United States and Canada by inter-library loan (ILL).

Articles from Doklady Akademii Nauk – Biological Sciences are available by ILL. Voprosy Ikhtiologii abstracts in English from 1996 to 2003 are available at the Voprosy Ikhtiologii website, as are full citations from the Doklady Akademii Nauk website.

Some articles indicated have abstracts available online from the Wild Salmon Center or Voprosy Ikhtiologii while others generally have sufficient citation information for locating a copy at a university library or via intra-library loan(ILL).

Peer-Reviewed Articles
  • Kuzishchin, K.V., Savvaitova, K.A., Gruzdeva M.A. 1999. Struktura cheshui kak kriterii differentsiatsii lokal’nykh populiatsii mikizhi Parasalmo mykiss iz rek zapadnoi Kamchatki i Severnoi Ameriki [Scale structure as criteria for the determination of local stocks of mikizha Parasalmo mykiss from western Kamchatkan and North American rivers] Voprosy Ikhtiologii [Journal of Ichthyology] 39(6):809-818.
  • Kuzishchin, K.V. and Savvaitova, K.A. 1998. Struktura cheshui mikizhi iz rek Severo-Zapadnoi Kamchatki i Sandy River (Alaska) [The Structure of steelhead scales from the rivers of Northwest Kamchatka and from the Sandy River of Alaska]. Voprosy ikhtiologii [Journal of Ichthyology] (in publication).The structure of scales is analyzed (number of annular rings, width of growth zones) in the river and marine years of life of the intrapopulation groups (typically anadromous, coastal, ‘half pounders’, freshwater river) and local stocks of Kamchatkan and American steelhead. The differences between neighboring populations in Kamchatka are not large. The population of the Sandy River differs from the Kamchatkan ones by the structure of the scale in the river period of life (more and wider annuli).
  • Kuzishchin, K.V., Pavlov, D.S., Savvaitova, K.A., Gruzdeva, M.A. and Pustovit, O.P. 2001. Downstream Migration of Juvenile Diadromous Kamchatka Trout Parasalmo mykiss in the Western Kamchatka Rivers. Voprosy ikhtiologii [Journal of Ichthyology] 41(3) 227-238. Abstract.
  • Savvaitova, K.A., Mina, M.V., Maksimov, V.A. 1975. Evolutionary Aspects of the Reproductive Ecology of Trout of the Genus Salmo in Kamchatka. Voprosy Ikhtiologii [Journal of Ichthyology] 15(1): 21-31.
  • Kuzishchin, K.V. and Savvaitova, K.A. 1996. Osteologicheskiye osobennosti tikhookeanskikh forelei (Salmo) Kamchatki v sviazi s opredeleniem ikh taksonomicheskogo statusa. [Osteological Peculiarities of Pacific Trouts
    (Genus Salmo) of Kamchatka in Connection with the Determination of their Taxonomic Status]     Voprosy ikhtiologii [Journal of Ichthyology] 36(4): 288-306. Abstract.
  • Mednikov, B.M., Shubina, Ye. A, Melnikova, M.N., Savvaitova, K.A. 1998. Problema rodovogo statusa tikhookeanskikh lososei i forelei. Genosistematicheskii analiz [The Problem of the Generic Status of Pacific Salmons and Trouts. A Genosystematic Analysis]. Voprosy ikhtiologii [Journal of Ichthyology] (in publication). Atlantic and Pacific salmons and trouts were studied by the method of electrophoretic analysis of nuclear DNA (taxonoprint). The DNA patterns of the species studied are divided into four groups: 1. Salmo salar; 2. Salmo trutta and closely related species; 3. Pacific trouts (including Kamchatkan forms); 4. Pacific salmons (O. masu, O. keta, O. gorbuscha, O. nerka, O. tshawytscha, and O. kisutsch). These groups correspond to their generic ranking. The merging of Parasalmo and Oncorhynchus into one genus Oncorhynchus is not confirmed at the genetic level and contradicts the international rules of zoological nomenclature.
  • Pavlov, D.S. and Savvaitova, K.A. 1996. Sovremennye problemy izucheniia i sokhraneniia vida Krasnoi knigi Rossii – kamchatskoi mikizhi [Current Problems of Study and Conservation of the Kamchatkan mikizha, a Species Listed in the Red Book of Russia] Voprosy ikhtiologii [Journal of Ichthyology] 36(8): 578-583.
    Abstract.
  • Pavlov, S.D. and Kuzishchin, K.V. 1999. Morfobiologicheskie i geneticheskie osobennosti mikizhi Parasalmo mykiss iz reki Voyampolki (zapadnaya Kamchatka) [Morphobiological and genetic pecularities of mikizha Parasalmo mykiss from Voyampolka River] Voprosy Ikhtiologii [Journal of Ichthyology] 39(6):748-760.
  • Pavlov, D.S., Savvaitova, K.A. and Kuzishchin, K.V. 1999. K probleme formirovaniia epigeneticheskikh variatsii zhiznennoi strategii u vida Krasnoiu knigi Rossii – kamchatskoi mikizhi Parasalmo mykiss (Salmonidae, Salmoniformes) [Regarding the problem of epigenetic variations of life-history strategy in the Kamchatkan Parasalmo (O.) mykiss, the species from Russian Red Data book] Doklady Akademii Nauk [Reports of the Russian Academy of Sciences – Biological Sciences] 367(5):709-713. (Russian)
  • Proebstel, D.S., Savvaitova, K.A., Kuzishchin, K.V. 1997. Native trout of Kamchatka: a glimpse into the past of North America // Wild Trout VI. Montana St. Univ. Bozeman. pp. 77-85.
  • Reshetnikov, Yu.S., Bogutskaya, N.G., Vasileva, E.D, Dorofeeva, Ye.A. Naseka, A.M., Popova, O.A., Savvaitova, K.A., Sidelava, V.G., and Sokolov, L.I. 1997. Spisok ryboobraznykh i ryb presnykh vod Rossii [List of the freshwater fishes of Russia] Voprosy ikhtiologii [Journal of Ichthyology] 37(6): 723-771.Gives a full list of 349 species of fishes living in the continental waters of Russia. Gives the names, the taxonomic changes, and additions (including data on the newly discovered trout in Kamchatka), defines the ranges of the species more
    precisely.
  • Savvaitova, K.A., Kuzishchin, K.V. and Maximov, S.V. 1999. Kamchatka steelhead: Population trends and life history variations, in Sustainable Fisheries Management: Pacific Salmon (Knudsen, E.E., ed.) Lewis Publishers, New York 1999: 195-206.
  • Savvaitova, K.A., Kuzishchin, K.V., Pavlov, D.S. 1999. Struktura lokal’nykh populiatsii mikizhi Parasalmo mykiss iz rek severo-zapadnoi Kamchatki i Severnoi Ameriki [The population structure of mikizha Parasalmo mykiss from Rivers of Northwest Kamchatka and North America] Voprosy Ikhtiologii [Journal of Ichthyology] 39(7): 504-516. Abstract.
  • Pavlov, D.S., Savvaitova, K.A., Kuzichshin, K.V. 2001. Theoretical aspects of the present-day distribution and forming of life history strategies in mikizha, Parasalmo mykiss (Salmonidae, Salmoniformes). Doklady Akademii Nauk [Reports of the Russian Academy of Sciences – Biological Sciences] Vol. 379: 344-347.At our period of time the range of mikizha in Asia is limited in Kamchatka, while throughout Kamchatkan waterbodies there is a strong difference in the ratio of phenotypes with different life history strategy. The most probable factors, determining occurrence and distribution of typically anadromous life history strategy pattern on Kamchatka can be considered the next ones: appearance of tundra rivers of tundra tributaries; appearance of deep pools of 2-5 m depth, that not freezing to the bottom during the winter to provide wintering of large non-feeding mature fish; appearance of the spawning ground, that fits for the spawning of large fish. The distribution of resident life history strategy is caused by the next reasons: appearance of the tundra rivers or tundra tributaries; appearance of non-freezing sections of the river with the spring waters, coming into the river bed, for wintering and feeding; appearance of the rivers with high gradient with transparent water, where fish is feeding in summer time; appearance of enough food and energetic resources in the river, for maturation not only males but females also.
  • Pavlov, D.S., Savvaitova, K.A., Kuzichshin, K.V. 2000. Dwarf Males and Reproductive Tactics in the Mikizha Parasalmo mykiss Walbaum (Salmonidae, Salmoniformes) from the Kamchatka Peninsula. Doklady Akademii Nauk [Reports of the Russian Academy of Sciences – Biological Sciences] Vol. 373: 382-385.
  • Savvaitova, K.A., Kuzishchin, K.V., Proebstel, D. 1998. Riady form forelei Zapadnoi Kamchatki. K probleme feneticheskogo raznobraziia gruppy [A Series of Forms of the Trout of Genus Salmo in Western Kamchatka. On the problem of the phenetic diversity of the group] Voprosy ikhtiologii [Journal of Ichthyology] 38(5): 366-373.
    Abstract.
  • Savvaitova, K.A. and Kuzishchin, K.V. 1998. Tendentsiia v izmenenii morfometricheskikh pokazatelei prokhodnoi formy Salmo mykiss iz rek severo-zapadnoi Kamchatki [Tendencies toward Change in the Morphometric Features of Diadromous mikizha Salmo mykiss from the Rivers of Northwest Kamchatka]. Voprosy ikhtiologii [Journal of Ichthyology] 38(3) 238- (in publication). Abstract.
  • Savvaitova, K.A. and Soverel, P. 1998. Discovered: Bering sea steelhead // International Journal of Salmon Conservation 1(1):2-3.
  • Savvaitova, K.A. and Kuzishchin, K.V. 1998. Struktura populiatsii mikizhi iz rek Kamchatki i Severnoi Ameriki [The Population Structure of mikizha from the rivers of Kamchatka and North America] Voprosy ikhtiologii [Journal of Ichthyology] (in publication).
  • Savvaitova, K.A., Kuzishchin, K.V., Maximov, S.V., and Pavlov, S.D. 1997. Struktura populiatsii kamchatskoi mikizhi v reki Utkholok (zapadnaya Kamchatka) [Population structure of mikizha, Salmo mykiss, in the Utkholok River (western Kamchatka)]. Voprosy ikhtiologii [Journal of Ichthyology]: 37(3) 216-225. Abstract.
  • Savvaitova, K.A., Kuzichshin, K.V., Maximov, S.V., Novikov, G.G. 1995. O raznoobrazii tikhookeanskikh forelei (rod Salmo) Kamchatki [On the diversity of Pacific trout (genus Salmo) in Kamchatka]. Voprosy Ikhtiologii [Journal of Ichthyology] 35(5):565-571.In 1994 an unusual trout was caught by anglers in the Tigil River basin. These specimens differed in a number of diagnostic characters from anadromous and freshwater forms of S. mykiss, inhabiting neighboring rivers. The trout from the Tigil River has several characters similar to those of S. clarki.
  • Soverel, P., Savvaitova, K.A., Kuzishchin, K.V., Maximov, S.V., and Pavlov, S.D. 1997. The Kamchatka steelhead project: a unique approach to understanding and conserving steelhead and trout of Kamchatkan peninsula, Russia // Wild Trout VI. Montana St. Univ. Bozeman. P. 39-44.Describes the 1996 WSC-MGU expedition on Kamchatka. The Kamchatka steelhead project expedition demonstrates a combination of scientific study of endangered species and protection of the unique resource by the elimination of poaching during the field expedition.
  • Stanford, J.A., Gayeski, N., Pavlov, D.S., Savvaitova, K.A. and Kuzishchin, K.V. 2002. Biophysical Complexity of the Krutogorova River (Kamchatka, Russia). Verh. Internat. Verein. Limnol. 28(3):1354-1361.The first examination of the biophysical complexity (geomorphology, benthic ecology, riparian structure) of a West
    Kamchatka steelhead river.
  • Pavlov, S.D., Kuzichsin, K.V. 1999. Morphobiological and Genetic Features of Kamchatka Steelhead Parasalmo mykiss from the Voyampolka River (Western Kamchatka). Voprosy Ikhtiologii [Journal of Ichthyology] 39(9): 713-725. Abstract.
  • Williams, R.N., Powell, M.S., Pavlov, S.D., and Proebstel, D.S. Forthcoming. Molecular Genetic Variation among rainbow trout Oncorhynchus mykiss Walbaum (Salmonidae, Salmoniformes) from the Kamchatka Peninsula. Submitted to Journal of Fish Biology.
  • Savvaitova, K.S. The Population Structure of Salmo mykiss in Kamchatka. 1975. Voprosy Ikhtiologii [Journal of Ichthyology] 15(6): 984-997.
  • Kuzichshin, K.V., Pavlov, S.D., Gruzdeva, M.A., Pavlov, D.S., Maksimov, S.V., Savvaitova, K.S. 2002 Spawning Population and Reproductive Ecology of the Freshwater Kamchatka Steelhead Parasalmo mykiss in the Basin of the Zhupanova River (East Kamchatka). Voprosy Ikhtiologii [Journal of Ichthyology] 42(8): 601-614. Abstract.
  • Savvaitova, K.S., Tutukov, M.A., Kuzichshin, K.V., Pavlov, D.S. 2002 Changes in the Population Structure of Mikizha Parasalmo mykiss from the Utkholok River, Kamchatka, during the Fluctuation in Its Abundance.
    Voprosy Ikhtiologii [Journal of Ichthyology] 42(3): 238-242. Abstract.
  • Kuzischin, K.V., Pustovit, O.P., Pavlov, D.S., Savvaitova K.A. 2002 Morphobiological Traits of Downstream-Migrating Juveniles of Parasalmo mykiss from Some Rivers of Western Kamchatka in Relation to Smolting. Voprosy Ikhtiologii [Journal of Ichthyology] 42(9): 720-732. Abstract.

Scientific Reports

Written for the Wild Salmon Center, Moscow State University, and the Ministry of Environment of the Russian Federation.

  • The Kamchatka Steelhead Project: Studies toward the preservation of a species in the Red Book of
    Russia), 1994 Science Report. (1995) 35 pp.
  • The Kamchatka Steelhead Project, 1995 Science Report (1996), 41 pp.
  • The Kamchatka Steelhead Project, 1996 Science Report(1997), 43 pp.
  • The Kamchatka Steelhead Project, 1997 Science Report (1998) 80 pp.
  • The Kamchatka Steelhead Project Five-Year Report, 1994-1998 (1999)..
  • The Kamchatka Steelhead Project, 1999 Science Report (2000) 108 pp. (Russian only).
  • The Kamchatka Steelhead Project, 2000 Science Report (2001) 108 pp. (Russian only).
  • The Kamchatka Steelhead Project, 2001 Science Report (2002) 87 pp. (Russian only).
  • The Kamchatka Steelhead Project, 2002 Science Report (2002) 93 pp. (Russian only).

Papers Presented

  • Pavlov, D.S., Savvaitova, K.A., and K.V. Kuzishchin. 1999. Epigeneticheskie variatsii zhiznennoi strategii mikizhi v rekakh zapadnoi Kamchatki i strategiia sokhraneniia vida Krasnoi knigi [Epigenetic variation of life history strategy of mikizha in western Kamchatkan rivers and strategy for conservation of Red data book species] // Proceedings of conference “The problems of protection and rational using of natural resources of Kamchatka.” Petropavlovsk-Kamchatsky, June 10-12, 1999: 81. (Russian only)
  • Savvaitova, K.A., Kuzishchin, K.V. 1997. The “series of forms” of Pacific trouts.
    To the question of the origin of the group     // Proceedings of the 1st Congress of Ichthyologists of Russia. Astrakhan’, September 1997: 51. (Russian only)
  • Savvaitova K.A., Kuzishchin K.V., Gruzdeva M.A., Pavlov D.S., Maksimov S.V., Pavlov S.D. 1999. Struktura lokal’nykh populiatsii mikizhi v rekakh zapadnoi Kamchatki i problemy ikh okhrany [The structure of local populations of mikizha in Western Kamchatkan Rivers and problems of their conservation] // Proceedings of conference “The problems of protection and rational use of natural resources of Kamchatka.” Petropavlovsk-Kamchatsky, June 10-12, 1999: 87-88. Russian only.
  • Savvaitova, K.A., Kuzishchin, K.V., Maksimov, S.V., Pavlov, S.D. 1997. Changes in population structure in anadromous steelhead from North-West Kamchatkan Rivers // Proceedings of the 1st Congress of Ichthyologists of Russia. Astrakhan, September 1997: 51-52. (Russian only)
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